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Invertebrates were identified to genus or species, with the exception of Nematoda , Turbellaria, and terrestrial invertebrates, which were typically identified only to Family or Order.
Furthermore they have several plesiomorphic characters, such as a myoepitheal pharyngeal bulb, comparable to the other Cycloneuralia, especially the Nematoda and Gastrotricha.
He discovered microscopic animals such as nematodes and rotifers.
For example, both the Nematoda and Platyhelminthes contain obligate parasites as well as free-living species.
Other ecdysozoan phyla include Nematoda , Tardigrada, and Onychophora; this classification thus removes onychophorans from the Arthropoda and makes them an outgroup.
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